At budbreak, and leaf senescence, the tree is expending a great deal of energy in order to:
In the former case, the tree is establishing the logistics for the 'food factories' for the growing season. It is moving a great deal of water, reserved carbohydrates and starches from the trunk and root system to the stems and twigs which will support the aforementioned food factories. The reserves will be there in the event the tree needs to establish neoformed leaf buds or take advantage of favorable growing conditions.
In the latter, the tree is actually withdrawing as much energy that it can from the spent leaves, and pulling whatever carbohydrate or starch reserve that it can from the twigs and stems, and downloading these excess reserves into the trunk and root system. Preformed buds are established and made ready for the following growing season.
So, if we prune, even a conservative amount, during these sensitive times (which generally only last 1-2 weeks per individual tree and vary by species), the tree may be robbed of the ability to transport and utilise excess reserves of starch, carbohydrates and water. Obviously, trees will be better able to handle the stress of a conservative pruning vs. a heavy prune. Proper abscission is necessary for the tree in order for it to remove what little energy it can from each leaf.
Am I wrong, or can someone explain it better?